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INFLUENCE OF SHORT-TERM VARIATION RAINFOREST DIVERSITY

By Rhett Butler   |  Last updated July 31, 2012
Rainforests and their diversity do not exist in a constant state, but are the product of a series of impacts including fires, tree falls, small-scale human clearing, and even lava flows. These events can increase forest diversity by giving new species a chance to grow in the absence of the towering canopy trees. The growth of new tree species spells new opportunities for their symbiotic species (for example new pollinators or seed dispersers).

Forests that are regularly stressed, like those affected seasonally by strong winds and storms, tend to be dwarfed with a less developed canopy and reduced diversity. "Typical" tall rainforests are typically found where they are protected from strong winds, as in valleys and certain geographical areas.

Within a relatively small area there can be great variations in forest dynamics. For example, in the terra firme rainforests of the Central Amazon—where average canopy tree age can exceed 300 years and some trees can be more than one thousand years old— forest turnover rates can be extremely low. In contrast, nearby floodplain forests may have turnover rates of less than 70 years due to migrating river channels that periodically undercut river banks and trees.

Diversity is usually sharply reduced in forests degraded by activities such as logging, burning, and agricultural development. Generally, when forest is logged, the dense canopy structure is disturbed, allowing more sunlight to penetrate to the forest floor. The forest is more likely to dry out, and less water can be recycled through the system of evaporation and transpiration. Many rainforest species are unable to cope with the changes in the forest microclimate and either move on or gradually perish. In addition, the loss of certain valuable hardwood trees to logging has a major impact on species with which they have interdependent relationships. Studies suggest that logging in any form reduces tropical forest diversity—studies around the world show declines of certain species, especially primates, birds, and insects in degraded forests. While there may be a local increase in the abundance and diversity of certain species, there is an overall regional or global decline in biodiversity due to the loss of species specially adapted to the conditions of undisturbed forest. Degraded forest is also more prone to be developed or burned by humans, severely reducing diversity. Heavy logging in the forests of Indonesia and Brazil was partly responsible for creating the dry forest conditions that drove the widespread forest fires of 1997-1998.


Fallen tree in Borneo
Fallen tree in Borneo. (Photo by R. Butler)



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  • Eldredge, N. and Gould S. ("Punctuated equilibrium: an alternative to phyletic gradualism." in T. Schopf, Models in Paleobiology, New York: WH Freeman 1972) introduce the idea of punctuated equilibrium as a new theory for evolution.
  • "Doomsday genes" which may enable species to undergo radical structural changes in mere generations in response to sudden environmental changes are discussed in Rutherford, S.L. and S. Lindquist, "HSP90 as a capacitor for morphological evolution," Nature 396: 336-342, 1998.